Epidemiology, Microbiology, Infectious Disease (Bovine, Companion Animal, Laboratory Animal, Primate, Ruminant, Wildlife) Zoonoses and public health; Epidemiology of zoonoses, foodborne zoonoses; Wildlife epidemiology and tropical medicine.

Epidemiology, zoonoses and veterinary public health Supervises MPVM student research projects

Plaque surveillance in wildlife and domestic animals for the State of California Zoonoses specialist to Calif. Dept. of Health Services Guest speaker to numerous organizations nationally and internationally Rabies epidemiology expert.

Co-Editor-in-Chief of Veterinary Research, 2002-present.

WHO Collaborating Center for New and Emerging Zoonoses